Glasgow Minimum Essential Medium was originally developed by lan MacPherson and Michael Stoker as a modification of Eagle's medium to study the genetic factors affecting cell competence. The polyoma virus was used to transform four fibroblast clones from a culture of baby hamster kidney cells. The medium was developed by modifying Eagle's BME by adding 10% tryptose phosphate and twice the normal concentration of amino acids and vitamins.
| Components shown as mg/liter |
| Calcium Chloride•2H2O | 264.92 |
| Ferric Nitrate•9H2O | 0.10 |
| Magnesium Sulfate | 97.67 |
| Potassium Chloride | 400.00 |
| Sodium Chloride | 6400.00 |
| Sodium Phosphate, anhy | 107.83 |
| L-Arginine•HCl | 42.00 |
| L-Cystine•2HCI | 31.29 |
| L-Glutamine | 292.00 |
| L-Histidine•HCl•H2O | 21.00 |
| L-Isoleucine | 52.40 |
| L-Leucine | 52.40 |
| L-Lysine•HCI | 73.10 |
| L-Methionine | 15.00 |
| L-Phenylalanine | 33.00 |
| L-Threonine | 47.60 |
| L-Tryptophan | 8.00 |
| L-Tyrosine•2Na•H2O | 52.19 |
| L-Valine | 46.80 |
| Choline Chloride | 2.00 |
| Folic Acid | 2.00 |
| myo-Inositol | 3.60 |
| Niacinamide | 2.00 |
| D-Pantothenic Acid•Ca | 2.00 |
| Pyridoxal•HCI | 2.00 |
| Riboflavin | 0.20 |
| Thiamine•HCI | 2.00 |
| D-Glucose | 4500.00 |
| Phenol Red, Sodium | 16.0 |
| Tryptose Phosphate Broth | 3000.00 |
| Total: 15.6g/L |
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